My first thought is skin infection - a bacterial infection or something. Could your vets (both OP and Lussy) test for that? I know I've seen mentions of hormone-caused quill loss (I think from Nancy), but I believe that case was a female hedgehog. Maybe Nancy will pop over here & have more to add. I'm not sure I have any more ideas, unfortunately. Poor hedgies!
~*~*~ Kelsey ~*~*~
RIP my sweet Lily ~ 6/12/08 - 1/20/12